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FLORA INDTCA.               '                      89
The ovaries of Anonace^ are generally very numerous and small, and closely
packed together. In Uvaria they are columnar., and quite straight, and grooved
along the inner face j but generally they are rounded on the hack, and oblong in shape.
They are usually very hairy, but sometimes perfectly glabrous. This character, though
constant in each species, is of no avail for the distinction of genera. The style is in-
variably terminal, and is either continuous with the ovary, and undistinguishable from
it except by the absence of a cavity, or separates by a joint. In the latter case the
mass of styles often coheres together by means of a viscid or gelatinous fluid. The
style is usually grooved OIL the inner face, and is stigmatic over its whole surface, and
often covered with papilli. Sometimes it is short and capitate, more generally ob-
long, and occasionally elongate and subulate. With occasional exceptions, which will
be noticed under the genera in which they occur, these characters seem constant.
The number of ovaries is of less value. In Xylopia, a very natural genus, they vary
from one to ten, and in Qrop/iea from three to fifteen. In A$inwia and other genera
their number is equally uncertain.
The number and position of the ovules are of great importance as generic charac-
ters. When solitary, the ovule is either erect from the base of the cell, as in Unona,
and Guatteria, or attached to the ventral suture, as in Ellipeia mid in some Miti-
vstff. Tu Artaliolrys aud Ana.ragorea there are always two collateral ovules, erect
from the base of the cell. \Yheu the ovules arc definite, aud attached to the ventral
suture, their number seems less constant. Thus, in Unona they vary from two to
eight, but are nearly constant in each species. In $rilini>a they vary from one to
two, and in Xylopia from two to six. In Pofyalt/tia and J?h<sa,nihu$ there are two
superposed ovules inserted very near the base of the cell, one of which seems occa-
sionally absent, in which case Polyatthiet is with difficulty distinguishable from
Chiatteria. When the ovules are numerous they arc arranged more or less distinctly
in two rows, and arc closely packed together: they are then occasionally subdcfinite,
especially where the ovary is very short, but this is hi no case a character of generic
value. The section Kentia of Malodorant, where they are reduced to two, is the only
very marked exception to the importance of the difference between definite aud inde-
finite ovules in the Order.
The shape of the stamens forms a very important character in A^onaceaft when-
ever it deviates from the ordinary type. This type, which dcpcuds mainly upon the
great compression of the anther, is nearly sessile, cuneate, tetragonal, with two
dorsal cells almost in contact with one another, nud the connective produced be-
yond the anthers into a depressed rounded head. More rarely the cells are distant,
and almost lateral. The process of the connective is, however, in some genera elon-
gated, and not at all depressed or truncate. lu one section of Uvaria, the anthers
are flat and almost foliaccous; and in the whole group of Saccopetatets they are ovoid,
with a scarcely conspicuous process of the connectivum. When the stamens-are
definite in number they are very irregular in shape, but usually trapezoidal, with a
thick fleshy connective and small dorsal anther-cells.
The torus varies remarkably in amount of development. Where the number of
ovaries and stamens is definite, it is very small; but in general it is large aud con-
spicuous, being sometimes cylindrical and elongated, as in llammcuhts or Magnolia,
but more generally conical, somewhat after the fashion of Jtubus, or broadly cylin-
drical and truncated. It is not unfrequently slightly concave iu the centre; and this
concavity becomes extreme in Xytopia, where the stamens are borne on the outside
of the torus, which completely encloses the ovaries: The modifications of this organ
are very constant, but not always sufficiently capable of definition to render them
available to the systcmatist.
The shape of the petals has been much neglected in the formation of genera.
Blume, however, has employed it as a sectional character in Ifaaria, under which
genus he has united most of the many-ovuled Anonacea, and also in Polyafthia,
in which he includes many of those with two ovules. The sections tlms formed
are highly natural, as the species included in them agree very1 closely iu habit_;
and we have accordingly raised them to the rank of genera, following an indica-